In a previous paper published in Science, they cited features like flat feet, dense bones and nostrils placed high on its head, as reasons behind claiming that Spinosaurus swam in waters.

Whales lack the bone structure and muscles to support their own weight ... Interestingly, theropod dinosaurs such as Spinosaurus and T. rex were also the only animals to exceed the largest land-based ...

Since its discovery in 1915, the biology and behaviour of the enormous Spinosaurus has puzzled palaeontologists worldwide. It was recently argued that the dinosaur was largely an aquatic predator, ...